When this trisomic male fly undergoes meiosis to form sperm, what genetic combin

The fruit fly Drosophila melanogaster has a haploid chromosome number of n = 4, and 2n = 8. Chromosome IV in this species is a tiny autosome. Flies trisomic for chromosome IV are fertile and have no apparent defects.

The eyeless (ey) and gawky (gw) loci are tightly linked on chromosome IV. Loss-of-function ey− and gw− alleles are recessive to ey and gw, respectively. Flies homozygous for ey− lack eyes, and flies homozygous for gw− have disrupted circadian rhythms.

Consider a male fly trisomic for chromosome IV, with each of the three chromosome copies bearing different allele combinations for these two loci: