Leonardo da Vinci propelled forward our understanding of science, particularly, anatomy. In what way ...

Leonardo da Vinci propelled forward our understanding of science, particularly, anatomy. In what way did he accomplish this?
 
  A) Leonardo hired prostitutes to pose for him so that he could study their bodies.
  B) Leonardo dissected pigs to gain a better understanding of musculature in the mammal.
  C) Leonardo dissected human bodies to gain a better understanding of the muscular-skeletal system.
  D) Leonardo was not a proponent of anatomical study.
  E) Leonardo paid people to stand, run, sit, etc., while he studied the movement of muscles.